0	10	Nonopsonic	nonopsonic	JJ
11	23	phagocytosis	phagocytosis	NN
24	26	of	of	IN
27	38	Pseudomonas	Pseudomonas	NNP
39	49	aeruginosa	aeruginosa	NN
50	52	by	by	IN
53	64	macrophages	macrophage	NNS
65	68	and	and	CC
69	86	polymorphonuclear	polymorphonuclear	JJ
87	97	leukocytes	leukocyte	NNS
98	106	requires	require	VBZ
107	110	the	the	DT
111	119	presence	presence	NN
120	122	of	of	IN
123	126	the	the	DT
127	136	bacterial	bacterial	JJ
137	146	flagellum	flagellum	NN
146	147	.	.	.

149	156	Whereas	whereas	IN
157	160	the	the	DT
161	170	mechanism	mechanism	NN
171	173	of	of	IN
174	184	nonopsonic	nonopsonic	JJ
185	197	phagocytosis	phagocytosis	NN
198	200	of	of	IN
201	212	Pseudomonas	Pseudomonas	NNP
213	223	aeruginosa	aeruginosa	NN
224	227	has	have	VBZ
228	232	been	be	VBN
233	242	described	describe	VBN
242	243	,	,	,
244	247	the	the	DT
248	257	bacterial	bacterial	JJ
258	265	ligands	ligand	NNS
266	274	required	require	VBN
275	278	are	be	VBP
279	285	poorly	poorly	RB
286	296	understood	understand	VBN
296	297	.	.	.

298	300	To	to	TO
301	309	identify	identify	VB
310	313	the	the	DT
314	323	requisite	requisite	JJ
324	333	bacterial	bacterial	JJ
334	341	ligands	ligand	NNS
341	342	,	,	,
343	350	studies	study	NNS
351	355	with	with	IN
356	364	isogenic	isogenic	JJ
365	372	mutants	mutant	NNS
373	375	of	of	IN
376	378	P.	P.	NNP
379	389	aeruginosa	aeruginosa	NNP
390	393	PAK	PAK	NNP
394	401	lacking	lack	VBG
402	406	pili	pili	NNS
406	407	,	,	,
408	416	flagella	flagella	NN
416	417	,	,	,
418	421	and	and	CC
422	425	the	the	DT
426	430	RpoN	RpoN	NNP
431	436	sigma	sigma	NN
437	443	factor	factor	NN
444	448	were	be	VBD
449	459	undertaken	undertake	VBN
459	460	.	.	.

461	464	The	the	DT
465	469	RpoN	rpon	NN
470	476	mutant	mutant	NN
476	477	,	,	,
478	485	lacking	lack	VBG
486	490	pili	pili	NNS
490	491	,	,	,
492	500	flagella	flagella	NN
500	501	,	,	,
502	505	and	and	CC
506	514	nonpilus	nonpilus	JJ
515	523	adhesins	adhesin	NNS
523	524	,	,	,
525	530	bound	bind	VBD
531	537	poorly	poorly	RB
538	541	and	and	CC
542	545	was	be	VBD
546	555	resistant	resistant	JJ
556	558	to	to	TO
559	568	ingestion	ingestion	NN
569	571	by	by	IN
572	576	both	both	CC
577	588	macrophages	macrophage	NNS
589	592	and	and	CC
593	604	neutrophils	neutrophil	NNS
604	605	.	.	.

606	610	Pili	Pili	NNP
611	615	were	be	VBD
616	619	not	not	RB
620	630	absolutely	absolutely	RB
631	639	required	require	VBN
640	643	for	for	IN
644	651	binding	binding	NN
652	654	or	or	CC
655	667	phagocytosis	phagocytosis	NN
668	670	of	of	IN
671	673	P.	P.	NNP
674	684	aeruginosa	aeruginosa	NNP
684	685	.	.	.

686	689	The	the	DT
690	698	presence	presence	NN
699	701	of	of	IN
702	703	a	a	DT
704	713	flagellum	flagellum	NN
714	717	was	be	VBD
718	721	not	not	RB
722	730	required	require	VBN
731	734	for	for	IN
735	742	binding	binding	NN
743	745	of	of	IN
746	748	P.	P.	NNP
749	759	aeruginosa	aeruginosa	NNP
760	762	to	to	TO
763	774	macrophages	macrophage	NNS
775	778	but	but	CC
779	782	was	be	VBD
783	791	critical	critical	JJ
792	795	for	for	IN
796	799	the	the	DT
800	810	subsequent	subsequent	JJ
811	826	internalization	internalization	NN
827	829	of	of	IN
830	833	the	the	DT
834	843	bacterium	bacterium	NN
843	844	,	,	,
845	855	suggesting	suggest	VBG
856	860	that	that	IN
861	865	this	this	DT
866	872	factor	factor	NN
873	875	or	or	CC
876	877	a	a	DT
878	885	surface	surface	NN
886	892	ligand	ligand	NN
893	903	associated	associate	VBN
904	908	with	with	IN
909	912	its	its	PRP$
913	921	assembly	assembly	NN
922	925	was	be	VBD
926	937	responsible	responsible	JJ
938	941	for	for	IN
942	953	stimulation	stimulation	NN
954	956	of	of	IN
957	967	nonopsonic	nonopsonic	JJ
968	980	phagocytosis	phagocytosis	NN
980	981	.	.	.